In Path of Exile 2, the Preserved Vertebrae is a unique Abyss-dropped currency item that plays a crucial role in endgame crafting. Specifically, it adds a desecrated modifier to a rare Waystone, an item type that functions as a powerful progression tool within the game's new systems.

Obtained from Abyss encounters. In Path of Exile 2, Abyss encounters drop a special line of currency items that desecrate rare gear. For weapons and quivers, this function comes through the Jawbone line of items.

Path of Exile 2 introduces unique bone-themed currency drops from Abyss encounters. Among them, the Gnawed Collarbone, Preserved Collarbone, and Ancient Collarbone are used to desecrate rare jewelry—specifically Amulets, Rings, and Belts. These items mirror the rib currency used on armour, but focus entirely on accessory slots.

Desecrated armour is one of the more intriguing crafting options in Path of Exile 2. It allows rare armour pieces to gain unique “desecrated modifiers” that sometimes can't be obtained through normal crafting. To start, you'll need special currency items dropped from Abyss encounters: Gnawed Rib, Preserved Rib, and Ancient Rib.

In Path of Exile 2, Desecrated modifiers play a special role in customizing your rare gear. To unlock these unique modifiers, players use Ancient Bones, a form of currency item tied to the Abyss system. These bones allow you to infuse rare items with desecrated affixes, opening the door to modifiers that might not normally appear. Ancient Bones are obtained primarily from Abyssal Troves, which spawn when you complete Abyss encounters, or inside the deeper layers of the Abyssal Depths. Keep in mind that when using these bones, if your rare item already has full affix slots, one of them will be randomly removed to make space for the desecrated modifier.

Desecrated modifiers can be applied to items using preserved bone currency. Bringing the item to the Well of Souls in Act 2 reveals the modifier, letting you choose one of three possible outcomes. While many options are standard modifiers, in endgame it's possible to encounter modifiers that normally cannot appear. Even before being revealed, a desecrated modifier occupies a prefix or suffix slot, and once revealed, it only rolls options of that type. Any item carrying a desecrated modifier displays a unique icon on its nameplate. Unrevealed modifiers appear as a line of green abyssal runes, while revealed exclusive modifiers show a faint green glow along the edges of the item's display.

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